Review of the Freshwater Mullets of Iran (Family Mugilidae)

Brian W COAD

Abstract

The systematics, morphology, distribution, biology, economic importance and conservation of the freshwater mullets of Iran are described, the species are illustrated, and a bibliography on these fishes in Iran is provided. Two species are found in the Caspian Sea basin as exotics (Chelon auratus, C. saliens), two species in southern Iranian fresh waters as native species (Mugil cephalus, Planiliza abu) and two species in brackish waters of rivers (Ellochelon vaigiensis, C. subviridis).

Keywords

Biology, Morphology, Mugilidae, Chelon, Ellochelon, Liza, Mugil, Planiliza.

Full Text:

PDF

References

Abbasi, K. 2006. Identification and distribution of fish fauna in Hevigh River (Guilan Province). Iranian Journal of Biology 18(4): 370-382. (In Farsi)

Abbasi, K.; Valipour, A.; Talebi Haghighi, D.; Sarpanah, A. & Nezami, Sh. 1999. Atlas of Iranian Fishes. Gilan Inland Waters. Gilan Fisheries Research Centre, Rasht. 113p. (In Farsi)

Abd, I.M.; Rubec, C. & Coad, B.W. 2009. Key biodiversity areas - Rapid assessment of fish fauna in southern Iraq, p. 161-171. In: Krupp, F., Musselman, J.L.; Kotb, M.M.A. & Weidig, I. (eds.), Environment, Biodiversity and Conservation in the Middle East, Proceedings of the First Middle Eastern Biodiversity Congress, Aqaba, Jordan, 20-23 October 2008. BioRisk 3(Special Issue), p. 219.

Abdoli, A. 2000. The Inland Water Fishes of Iran. Iranian Museum of Nature and Wildlife, Tehran. 378p. (In Farsi)

Abdoli, A. & Naderi, M. 2009. Biodiversity of Fishes of the Southern Basin of the Caspian Sea. Abzian Scientific Publications, Tehran. 243 p. (In Farsi)

Abdoli, A.; Naderi, M.; Aboo, M.; Fazli, H. & Afraee, M. A. 1996. Spawning time and fecundity of the mullet, Liza auratus (sic) in south eastern part of the Caspian Sea. Abzeeyan, Tehran 7(2): 24-26. (In Farsi)

Abdolmalaki, Sh. 2001. Catch statistic review of mullet fishery in Iranian coastal waters of the Caspian Sea. Iranian Journal of Fisheries Sciences 9(4): 39-52. (In Farsi)

Abdolmalaki, Sh.; Amirkhani, A.; Borani, M.; Ghaninejad, D.; Rastin, R.; Porgholami, A. & Moradkhah, S. 1998. A survey on catch status and sexual maturity of mullets in Iranian coastal water of the Caspian Sea (Gilan Province) in October 1998. Research Report, Fisheries Research Center of Gilan Province, Bandar Anzali. 13p. (In Farsi)

Abdolmalaki, Sh. & Psuty, I. 2007. The effects of stock enhancement of pikeperch (Sander lucioperca) in Iranian coastal waters of the Caspian Sea. ICES Journal of Marine Sciences 64(5): 973-980.

Agah, H.; Leermakers, M.; Elskens, M.; Fatemi, S.M.R. & Baeyens, W. 2007. Total mercury and methyl mercury concentrations in fish from the Persian Gulf and the Caspian Sea. Water, Air, & Soil Pollution 181(1-4): 95-105.

Aghili, K. & Mohammadi, F. 2011. Study on catching trend of Liza saliens in the Gorgan Bay. Journal of Biology Science 4(4): 47-56.

Ahmad, T.; Slaman (sic), N.A. & Hussain, N.A. 1983. Effects of some experimental conditions on the behaviour and survival of Liza abu (Heckel) from Basrah, Iraq. Journal of the Faculty of Marine Science, Jeddah 3: 111-117.

Ahmad, T.A. & Hussain, N.A. 1982. Observations on the food of young Liza abu (Heckel) from Salihyia River (Basrah, Iraq). Iraqi Journal of Marine Science, Basrah 1(1): 79-88.

Ahmad, T.A.; Salman, N.A. & Hussain, N.A. 1985. Effect of some laboratory controlled conditions on the survival of Liza abu (Heckel). Kuwait Bulletin of Marine Science 1985(6): 125-131.

Ahmed, S.M. & Hussain, N.A. 2000. Egg and larvae of mullets (Mugilidae) in the north western Persian Gulf. Pakistan Journal of Marine Biology 6(1): 1-7.

Ahmed, S.M. & Hussain, N.A. 2003. Spawning sites and migration routes of Liza subviridis and Liza carinata larvae in Iraqi marine waters. Marina Mesopotamica 18(2): 105-111.

Ahmed, S.M.; Salman, N.A. & Hashim, A.A. 2002. Osmotic regulation in Liza abu and Liza carinata (Pisces: Mugilidae) from estuaries of southern Iraq. Marina Mesopotamica 17(2): 293-304.

Akbari, H. 1998. Mullets found in Hormozgan moshtas in the Persian Gulf. Iranian Journal of Fisheries Sciences 7(3): 1-12, 1. (In Farsi)

Akbari, H. 2002. Frequency of Mugil cephalus in catch composition of stake-nets in the Hormozgan Province waters. Iranian Journal of Fisheries Sciences 11(1): 1-8. (In Farsi)

Akbary, P.; Arbabi, K.; Blochzehi, M.; Jamishzehi, H. & Tahmasebi, S. 2015. Effect of garlic extract on growth, feed and carcass composition of Mugil cephalus. Journal of Fisheries (Iranian Journal of Natural Resources) 68(3): 343-353.

Akhondzadeh Basti, A.; Misaghi, A.; Zahraei Salehi, T. & Kamkar, A. 2006. Bacterial pathogens in fresh, smoked and salted Iranian fish. Food Control 17(3): 183-188.

Al-Daham, N.K. & Wahab, N.K. 1991. Age, growth and reproduction of the greenback mullet, Liza subviridis (Valenciennes), in an estuary in southern Iraq. Journal of Fish Biology 38(1): 81-88.

Al-Daham, N.K. & Yousif, A.Y. 1990. Composition, seasonality and abundance of fishes in the Shatt Al-Basrah Canal, an estuary in Southern Iraq. Estuarine, Coastal and Shelf Science 31(4): 411-421.

Al-Hassan, L.A.J. 1984. Meristic comparison of Liza abu from Basrah, Iraq and Karkhah River, Arabistan, Iran. Cybium 8(3): 107-108.

Al-Hassan, L.A.J. 1987. Vertebral anomalies in fishes collected from Iraq and Arabistan, Iran. Pakistan Journal of Zoology 19(1): 95-97.

Al-Hassan, L.A.J. & Al-Seyab, A.A. 1992. A record size mugilid fish, Liza subviridis (Valenciennes, 1836) from the marsh area, southern part of Iraq. Pakistan Journal of Zoology 24(4): 354-355.

Al-Hassan, L.A.J.; Hussain, N.A. & Soud, K.D. 1989. A preliminary, annotated check-list of the fishes of Shatt Al-Arab River, Basrah, Iraq. Polskie Archiwum Hydrobiologii 36(2): 283-288.

Al-Hassan, L.A.J. & Madhi, A.A. 1987. Enzyme polymorphisms in the mullet, Liza dussumeiri (sic) from Shatt al-Arab River, Khor al-Zubair and the Persian Gulf. Biochemical Systematics and Ecology 15(2): 269-271.

Al-Hassan, L.A.J. & Na’ama, A.K. 1986. Some vertebral abnormalities in fishes collected from the Euphrates in Iraq. Indian Journal of Fisheries 33(2): 237-241.

Al-Hisnawi, F. 1990. Biology of the greenback grey mullet Liza subviridis (Val., 1836) in Khor Al-Zubair, south of Iraq. M.Sc. Thesis, University of Basrah. 116p. (In Arabic)

Ali, A.S.K.; Balasem, A.N. & Mutar, A.J. 1999. A preliminary study on the sensitivity of Liza abu (Heckel) to arsenic in comparison with eight species of local and breeding fishes. The Veterinarian 9(3): 35-42.

Ali, M.; Hedayatifard, M.; Pourgholam, R.; Farahani, Sh. & Rezaie, M. 2011. The study of brining and vacuum packaging on some quality aspects such as lipid deterioration of Liza aurata stored at 4ºC. Journal of Animal Biology 3(2): 25-31.

Ali, T.S.; Hantoush, A.A. & Jabir, A.A. 2004. Nutritional value of some Iraqi marine fishes. Marina Mesopotamica 19(2): 199-209.

Ali, T.S.; Hussain, N.A. & Younis, K.H. 1999. Stock assessment of greenback grey mullet Liza subviridis (Valenciennes, 1836) in Khor Abdullah, northwest Persian Gulf. Marina Mesopotamica 14(2): 394-407.

Al-Nasiri, S.K. & Hoda, S.M.S. 1975. Survey of fish fauna of Shatt-Al-Arab (from Abu-al-Khasib to Karmat Ali). Bulletin of the Basrah Natural History Museum 2: 36-46.

Al-Nasiri, S.K. & Hoda, S.M.S. 1976. A guide to the freshwater fishes of Iraq. Basrah Natural History Museum Publication. 124p.

Al-Nasiri, S.K. & Islam, A.K.M.S. 1978. Age and growth of Liza abu (Heckel) (Perciformes: Mugilidae) from Southern Iraq. Bangladesh Journal of Zoology 6(2): 85-89.

Al-Nasiri, S.K., Sarker, A.L. & Hoda, S.M.S. 1977. Feeding ecology of a mugilid fish, Liza abu (Heckel) in Basrah, Iraq. Bulletin of the Basrah Natural History Museum 4: 27-40.

Al-Nasiri, S.K. & Sirajul Islam, A.K.M. 1978. Age and growth of Liza abu (Heckel) (Perciformes: Mugilidae) from Southern Iraq. Bangladesh Journal of Zoology 6(2): 85-89.

Al-Shamma'a, A.A. & Jasim, Z.M. 1993. The natural food of Liza abu during the flood in Al-Hammar Marsh, South Iraq. Zoology in the Middle East 9: 59-64.

Al-Shamma'a, A.A.; Mohammad, M.A.; Shalash, F.J. & Nashaat, M.R. 2009. Seasonal variation in the natural diet of khisni Liza abu (Heckel, 1843) from Tigris River, Iraq. Alanbar University Journal for Pure Science 3(3): 66-73. (In Arabic).

Al–Shamma’a, A.A.; Mohammad, M.A. & Thykum, M. A–R. 1995. The natural food of Liza abu (Heckel, 1843), from Tigris River near the lower reaches of Diyala River Al–Za’afaraniya, Baghdad. Marina Mesopotamica 8(2)(1993): 356–365. (In Arabic)

Al-Yamour, K.Y.; Allose, S.B. & Al-Jafary, A.R. 1988. Some biological aspects of Liza abu (Heckel) Mugilidae, in Al-Daoodi Drain. (Baghdad, Iraq). Journal of Environmental Science and Health, Part A, Environmental Science and Engineering 23(5): 497-508.

Amini, F. 1989. Biology of mullet species and their adaptation to fresh water. In: Proceedings of the National Conference on Fisheries Resource Management of the Caspian Sea, Babolsar, Iran, p. 73-79. (In Farsi)

Amini Ranjbar, Gh. & Sotoudebnia, F. 2005. Investigation of heavy metals accumulation in muscle tissue of Mugil auratus in relation to standard length, weight, age and sex. Iranian Journal of Fisheries Sciences 14(3): 1-18. (In Farsi)

Amiri, K.; Bani, A.; Abd Almaleki, Sh. & Alijanpour, N. 2012. A study on reproductive investment of leaping grey mullet (Liza saliens) in the southwest Caspian SFea (Bandar Anzali). Journal of Fisheries (Iranian Journal of Natural Resources) 65(3): 235-242.

Ardalan, A.; Sharif, A.A. & Farahani, Z. 2012. Trace elements accumulation and nutritive value in muscle tissue of golden mullet from the Kiashahr Port (Caspian Sea). Advances in Environmental Biology 6(1): 232-235.

Askary Sary, A. & Beheshti, M. 2012. Cadmium, iron, lead and mercury bioaccumulation in abu mullet, Liza abu, different tissues from Karoun and Karkheh rivers, Khozestan, Iran. Bulletin of Environmental Contamination and Toxicology 88(2): 158-161.

Askary Sary, A. & Velayatzadeh, M. 2013. Bioaccumula-tion lead and zinc metals in the liver and muscle of Cyprinus carpio, Rutilus frsii kuttom (sic) and Liza auratus. Journal of Food Hygiene 3(1)(9): 89-99.

Askary Sary, A.; Velayatzadeh, M. & Beheshti, M. 2012. Determination of heavy metals in Liza abu from Karkheh and Bahmanshir Rivers in Khoozestan from Iran. Advances in Environmental Biology 6(2): 579-582.

Aslan Parviz, H. 1991. Mullet fish of the Caspian Sea. Abzeeyan 14: 20-25. (In Farsi)

Avanesov, E.M. 1972. Present spawning conditions of mullets (genus Mugil) in the Caspian Sea. Journal of Ichthyology 12(3): 419-425.

Azari Takami, G.; Amini, F.; Ranjbar, T. & Khodaii, A. 1997. Experimental survey of Caspian Sea mullets culture in brackish water of salt desert region of Iran. Journal of the Faculty of Veterinary Medicine, University of Tehran 51(1-2): 59-69. (In Farsi)

Bagheri, A.; Kamrani, E. & Esmaeili, H.R. 2010. Identified fishes of Kol River in Hormuzgan Province. Iranian Scientific Fisheries Journal 19(2): 143-148. (In Farsi)

Bagnis, R.; Berglund, F.; Elias, P.S.; van Esch, G.J.; Halstead, B.W. & Kojima, K. 1970. Problems of toxicants in marine food products. 1. Marine biotoxins. Bulletin of the World Health Organization 42: 69-88.

Balasem, A.N.; Ali, A-S.K. & Mutar, A.J. 2002. Sensitivity of some freshwater fishes for water pollutants. Proceedings of International Symposium on Environmental Pollution Control and Waste Management, (EPCOWM’2002), 7-10 January 2002, Tunis, p. 38-42.

Baltz, D.M. 1991. Introduced fishes in marine systems and inland seas. Biological Conservation 56(2): 151-177.

Bamshad, M.; Teimori, A.; Askari, M. & Majdzadeh, S.M. 2015. Morphology of the saccular otolith in Liza aurata (Risso, 1810) from coastal habitats of Caspian Sea. The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Banagar, G.R.; Kiabi, B.H.; Homayoonnezhad, I.; Piri, I. & Amirian, P. 2008. Biodiversity of fish species in Haraz River (an ecological approach). World Applied Sciences Journal 5(1): 5-11.

Barati, A. 2009. Diet and growth of chicks of the Great Cormorant, Phalacrocorax carbo, at Ramsar, northern Iran. Zoology in the Middle East 46(1): 29-36.

Barzegar, M. & Jalali, B. 2009. Crustacean parasites of fresh and brackish (Caspian Sea) water fishes of Iran. Journal of Agricultural Science and Technology 11: 161-171.

Barzegar, M.; Raeisi, M.; Bozorgnia, A. & Jalali, B. 2008. Parasites of eyes of fresh and brackish water fishes in Iran. Iranian Journal of Veterinary Research, Shiraz University 9(3)(24): 256-261.

Beheshti, M.; Askarysary, A.; Khodadadi, M. & Velayatzadeh, M. 2011. Measurement and comparison of heavy metals (Cu, Fe, Zn, Mn) in muscle, liver and gill organs of a special fish (Liza abu) in the Dez River in Khusestan Province. Journal of Wetland Ecobiology 2(6): 71-79. (In Farsi)

Behrooz, M.; Norouzi, M. & Nazemi, A. 2014. Genetic structure of golden mullet (Liza aurata) in the south Caspian Sea (Mazandaran Province) using microsatellite markers. Breeding and Aquaculture Sciences Quarterly 1(1): 11-16

Behrouzirad, B.; Rasekh, A. & Navidi, M. 2013. An estimation of consumption amount of fish by Great Cormorant (Phalacrocorax carbo) on fish culture pools in Shooshtar area (Khouzestan). Journal of Wetland Ecobiology 5(3): 17-30. (In Farsi)

Behruz, M.; Norouzi, M.; Sameie, M.H. & Zandavar, H. 2015. Genetic variation of golden mullet (Liza aurata) in Gomishan and Anzali wetlands. The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Berg, L.S. 1949. Presnovodnye ryby Irana i sopredel'nykh stran [Freshwater fishes of Iran and adjacent countries]. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 8: 783-858.

Berra, T.M. 2001. Freshwater Fish Distribution. Academic Press, San Diego. 604p.

Binaii, M.; Ghiasi, M.; Pourgholam, R.; Zorriehzahra, J.; Saidi, A.A. & Behrozi, S. 2010. Comparative evaluation of hematological factors of healthy and suspected to viral nervous necrosis mullets in Mazandaran province. In: Proceedings of the 16th Iranian Veterinary Congress, Tehran, Iran. pp. 200-201 (In Farsi).

Blegvad, H. & Løppenthin, B. 1944. Fishes of the Iranian Gulf. Einar Munksgaard, Copenhagen. 247 pp., XII pls. (1999 translation into Farsi by E. Etemad & B. Mokayyer with supplement, Tehran University Publications No. 1744: 26 + 416p.).

Bykhovski, B.E. 1949. Monogeneticheskie Sosal'shchiki Nekotorykh Ryb Irana, Sobrannye Akad. E.N. Pavlovskim [Monogenetic trematodes of some fishes in Iran, collected by Acad. E.N. Pavlovski]. Trudy Zoologicheskogo Instituta Akademii Nauk SSR 8: 870-878.

Caspian Environmental Programme. 1998. Caspian Sea Environment National Report, Islamic Republic of Iran. Baku, Azerbaijan. Pagination various.

Çelik, U.; Altınelataman, C.; Dinçer, T. & Acarlı, D. 2012. Comparison of fresh and dried flathead grey mullet (Mugil cephalus, Linnaeus 1758) caviar by means of proximate composition and quality changes during refrigerated storage at 4±2ºC. Turkish Journal of Fisheries and Aquatic Sciences 12(1): 1-5.

Chan, E.H. & Chua, T.E. 1979. The food and feeding habits of greenback grey mullet, Liza subviridis (Valenciennes), from different habitats and at various stages of growth. Journal of Fish Biology 15(2): 165-171.

Chan, E.H. & Chua, T.E. 1980. Reproduction in the greenback grey mullet, Liza subviridis (Valenciennes, 1836). Journal of Fish Biology 16(5): 505-519.

Chelemal M.; Jamili S. & Sharifpour, I. 2009. Reproductive biology and histological studies in abu mullet, Liza abu in the water of the Khozestan Province. Journal of Fisheries and Aquatic Science 4(1): 1-11.

Chelemal Dezfoul Nejad, M. 2007. Study on biological characteristics, sexual maturation and reproduction of Liza abu in the water of Khozestan Province. Ph.D. Thesis, Islamic Azad University, Tehran. 159p. (In Farsi)

Coad, B.W. 1979. Poisonous and venomous freshwater fishes of Iran. Pahlavi Medical Journal 9(4): 388-407.

Coad, B.W. 1987. Zoogeography of the Freshwater Fishes of Iran, pp. 213-228. In: Krupp, F., Schneider, W. & Kinzelbach, R. (eds.). Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz, 1985. Beihefte zum Tübinger Atlas des Vorderen Orients, Reihe A (Naturwissenschaften), 28, Dr. Ludwig Reichert Verlag, Wiesbaden, 338p.

Coad, B.W. 1991. Fishes of the Tigris-Euphrates basin: A Critical Checklist. Syllogeus, Ottawa 68: 1-49.

Coad, B.W. 1996. Exotic and transplanted fishes in Southwest Asia. Publicaciones Especiales Instituto Español de Oceanografía 21: 81-106.

Coad, B.W. 1997. Freshwater fishes of Iranian and Pakistani Baluchistan. Biologia, Lahore, 42(1&2) (1996): 1-18.

Coad, B.W. 1998. Systematic biodiversity in the freshwater fishes of Iran. Italian Journal of Zoology, 65 (Supplement): 101-108. (Proceedings of the Ninth Congress of European Ichthyologists (CEI-9) "Fish Biodiversity" organised in Naples at the University Federico II and held in Trieste - Italy, 24-30 August 1997).

Coad, B.W. 2010. Freshwater Fishes of Iraq. Pensoft Publishers, Sofia-Moscow. Pensoft Series Faunistica, 93. 294p.

Coad, B.W. & Abdoli, A. 1993. Exotic fish species in the fresh waters of Iran. Zoology in the Middle East 9: 65-80.

Coad, B.W. & Abdoli, A. 1996. Biodiversity of Iranian freshwater fishes. Abzeeyan, Tehran 7(1): 4-10. (In Farsi)

Crosetti, D. & Blaber, S. 2016. Biology, Ecology and Culture of Grey Mullet (Mugilidae). CRC Press, Taylor & Francis, Boca Raton. 529p.

Daoud, H.A.M.; Abid, W.B. & Al-Doori, T.Y. 1998. Observations on the reproductive biology of Liza abu (Heckel) from Tigris River. The Veterinarian 8(1): 154-164.

Daryanabard, G.R.; Shabani, A.; Kaymaram, F. & Gorgin, S. 2009. Reproduction and maturity of golden grey mullet (Liza aurata Risso, 1810) in Iranian waters of the Caspian Sea. Journal of Agricultural Sciences and Natural Resources 16(Special Issue 2): 77-88.

Deĝer, D.; Ünlů, E. & Gaffaroĝlu, M. 2013. Karyotype of mullet Liza abu Heckel, 1846 (Pisces, Mugilidae) from the Tigris River, Turkey. Journal of Applied Ichthyology 29(1): 234-236.

Dmitriev, N.A. 1946. Kefal' v iranskikh vodakh Kaspiya [Mullet in the Iranian waters of the Caspian]. Priroda 1946(12): 74-75.

Durand, J.-D.; & Borsa, P. 2015. Mitochondrial phylogeny of grey mullets (Acanthopterygii: Mugilidae) suggests high proportion of cryptic species. Comptes Rendus Biologies 338(4): 266-277.

Durand, J-D.; Chen, W-J.; Shen, K-N.; Fu, C. & Borsa, P. 2012. Genus-level taxonomic changes implied by the mitochondrial phylogeny of grey mullets (Teleostei: Mugilidae). Comptes Rendus Biologies 335: 687-697.

Durand, J.-D.; Shen, K.-N.; Chen, W.-J., Jamandre, B.W.; Biel, H.; Diop, K.; Nirchio, M.; Garcia de León, F.J.; Whitfield, A.K.; Chang, C.-W. & Borsa, P. 2012. Systematics of the grey mullets (Teleostei: Mugiliformes: Mugilidae): Molecular phylogenetic evidence challenges two centuries of morphology-based taxonomy. Molecular Phylogenetics and Evolution 64(1): 63-92.

Emadi, H. 1993. The status of aquaculture in Iran. Abzeeyan, Tehran 4(2): VII-IIX. In English.

Epler, P.; Bartel, R.; Szczerbowski, J.A. & Syzpuła, J. 2001. The ichthyofauna of lakes Habbaniya, Tharthar and Razzazah. Archiwum Rybactwa Polskiego 9(supplement 1): 171-184.

Epler, P.; Sokołowska-Mikołajczyk, M.; Popek, W.; Bieniarz, K.; Kime, D.E. & Bartel, R. 1996. Gonadal development and spawning of Barbus sharpei (sic), Barbus luteus and Mugil hishni in fresh and saltwater lakes of Iraq. Archiwum Rybactwa Polskiego 4(1): 113-124.

Eschmeyer, W.N.; Fricke, R. & van der Laan, R. 2016. Catalog of Fishes: genera, species, references (http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp).

Eskandari, G.; Sabz Alizadeh, S.; Dehgan Madiseh, S. & Mayahi, Y. 2007. Fish population’s structures in the Dez Dam Lake. Pajouhesh va Sazandegi 19(1): 123-129. (In Farsi)

Eslami, A.H. & Anwar, M. 1971. Occurrence and intensity of the infestation by Caryophyllaeus fimbriceps in carp and mullet (new host) in Iran. Rivista italiana di Piscicoltura e Ittiopatologia 6(1): 21-24.

Esmaeili, H.R.; Babai, S.; Gholamifard, A.; Pazira, A.; Gholamhosseini, A. & Coad, B.W. 2015. Fishes of the Persis region of Iran: an updated checklist and ichthyogeography. Iranian Journal of Ichthyology 2(3): 201-223.

Esmaeili, H.R.; Coad, B.W., Gholamifard, A.; Nazari, N. & Teimory, A. 2011. Annotated checklist of the freshwater fishes of Iran. Zoosystematica Rossica 19(2): 361-386.

Esmaeili, H.R.; Coad, B.W.; Mehraban, H.R.; Masoudi, M.; Khaefi, R.; Abbasi, K.; Mostafavi, H. & Vatandoust, S. 2014. An updated checklist of fishes of the Caspian Sea basin of Iran with a note on their zoogeography. Iranian Journal of Ichthyology 1(3): 152-184.

Esmaeili, H.R.; Khaefi, R.; Sayyadzadeh, G.; Tahami, M. S.; Parsi, B. & Gholamifard, A. 2014. Scale surface microstructure and scale size in three mugilid fishes (Teleostei, Mugilidae) of Iran from three different habitats. Istanbul University Faculty of Science Journalof Biology 73(1): 31-42.

Esmaeili, H.R.; Teimori, A.; Owfi, F.; Abbasi, K. & Coad, B.W. 2014. Alien and invasive freshwater fish species in Iran: Diversity, environmental impacts and management. Iranian Journal of Ichthyology 1(2): 61-72.

Esmaeili, H.R.; Mehraban, H.; Abbasi, K.; Keivany, Y. & Coad, B. 2017. Review and updated checklist of freshwater fishes of Iran: Taxonomy, distribution and conservation status. Iranian Journal of Ichthyology 4(Suppl. 1): 1-114.

Faddagh, M.S.; Ahmed, S.M. & Husain, S.A. 2004. Impact of gas oil on innic (sic) regulation, chloride cells and histopathological alterations in gills of mugilid Liza abu (Heckel) juveniles. Marina Mesopotamica 19(1): 36-52.

Farahnak, A. 2000. Fish Anisakidae in Khuzestan province of South West Iran. Bulletin of the Scandinavian Society for Parasitology 10(2): 71.

Farahnak, A.; Mobedi, I. & Tabibi, R. 2002. Fish Anisakidae helminthes in Khuzestan Province, south west of Iran. Iranian Journal of Public Health 31(3-4): 129-132.

Farid-Pak, F. No date. Société anonyme des Pècheries (sic) iraniennes. Bank Melli Iran Press, Tehran. 10p.

Fazeli, M.S.; Abtahi, B. & Sabbagh Kashani, A. 2005. Assessing Pb, Ni and Zn accumulation in the tissues of Liza aurata in the south Caspian Sea. Iranian Journal of Fisheries Sciences 14(1): 65-78. (In Farsi)

Fazli, H. 1998. Some biological characteristics of Liza aurata in the southern coasts of the Caspian Sea. Iranian Journal of Fisheries Sciences 7(3): 41-56. (In Farsi)

Fazli, H. 2000. Study on some biological characteristics of Liza saliens in southern part of the Caspian Sea. Iranian Journal of Fisheries Sciences 8(4): 29-42. (In Farsi)

Fazli, H. 2014. Species composition of bonyfish in the Iranian waters of the Caspian Sea during 1927-2013. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Fazli, H.; Daryanabard, G.H., Abdolmaleki, S. & Bandani, G.A. 2013. Stock assessment and management implications of golden grey mullet (Liza aurata Risso, 1810) in Iranian waters of the Caspian Sea. Journal of Applied Ichthyology, 29(2): 431-436.

Fazli, H. & Daryanabard, G.R. 2012. Qualitative assessment of golden grey mullet stocks (Liza aurata Risso, 1810) in the Caspian Sea (1991-2011). Journal of Fisheries (Iranian Journal of Natural Resources) 65(3): 307-315.

Fazli, H. & Ghaninejad, D. 2004. Some biological aspects and catch trend of mullets in the southern Caspian Sea. Iranian Journal of Fisheries Sciences 13(1): 97-114. (In Farsi)

Fazli, H.; Ghaninejad, D.; Janbaz, A.A. & Daryanabard, R. 2008. Population ecology parameters and biomass of golden grey mullet (Liza aurata) in Iranian waters of the Caspian Sea. Fisheries Research 93: 222-228.

Fazli, H.; Janbaz, A.A.; Taleshian, H. & Bagherzadeh, F. 2008. Maturity and fecundity of golden grey mullet (Liza aurata Risso, 1810) in Iranian waters of the Caspian Sea. Journal of Applied Ichthyology 24(5): 610-613.

Fazli, H.; Kor, D.; Daryanabard, Gh. & Tavakoli, M. 2014. Distribution and diversity of fishes in bottom trawls in Iranian part the Caspian Sea. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Fazli, H.; Kor, D.; Daryanabard, G.R.; & Tavakoli, M. 2014. Spatial and temporal distribution of mullets in Iranian waters of the Caspian Sea. Journal of Fisheries 8(2): 53-62.

Fereidouni, M.S.; Akbary, P. & Soltanian, S. 2015. Survival rate and biochemical parameters in Mugil cephalus (Linnaeus, 1758) larvae fed garlic (Allium sativum L.) extract. American Journal of Molecular Biology 5: 7-15.

Fischer, W. & Bianchi, G. 1984. FAO Species Identification Sheets for Fishery Purposes. Western Indian Ocean (Fishing Area 51). Food and Agriculture Organization, Rome. 6 volumes.

Ghadirnejad, H. 1996. Population dynamics of grey mullet species (Liza aurata and L. saliens) in the southern Caspian Sea. Ph.D. Thesis, University of Wales, Swansea. 207 pp.

Ghadirnejad, H. & Ryland, J. S. 1996. A study of food and feeding of grey mullets in the southern Caspian Sea, p. 137-144. In: GUTSHOP '96, Feeding ecology and nutrition in fish: symposium and proceedings, American Fisheries Society, Physiology Section, Bethesda, Md. 214p.

Ghalichi, A. 2003. The survey of sexual maturity in Mugil cephalus so measuring sex hormones and histologic studies. Ph.D. Thesis, Islamic Azad University, Tehran. 155p. (In Farsi)

Ghaneh, R. 2011. Study of genetic diversity of Liza saliens in the southern coasts of the Caspian Sea by mtDNA method. M.Sc. Thesis, Islamic Azad University, Iran. (In Farsi)

Ghaninejad, D. 2011. Maturity stages, gonado-somatic index (GSI) and fecundity of leaping grey mullet, Liza saliens (Risso, 1810) in the western part of Iranian waters of the Caspian Sea (Guilan, Province, Iran). Asian Fisheries Science 24(2): 115-226.

Ghaninejad, D.; Abdolmalaki, S. & Kulyev, Z. M. 2010. Reproductive biology of the golden grey mullet, Liza aurata in the Iranian coastal waters of the Caspian Sea. Iranian Journal of Fisheries Sciences 9(3): 402-411.

Ghaninejad, D.; Parafkandeh Haghighi, F.; Abdolmaleki, Sh.; Nahrevar, R.; Khedmati, K.; Rastin, R. & Nikpour, M. 2012. Morphometric and meristic characteristics of Liza aurata Risso 1810 in the south of Caspian Sea. Journal of Fisheries 6(2): 31-43.

Ghasemi, M.; Zorriehzahra, M.J.; Sharifpour, E. & Haghighi Karsidani, S. 2013. Detection of betanodavirus antigen associated with viral nervous necrosis (VNN) in tissue sections of apparently healthy golden grey mullets, Liza auratus, by histopathology examination and indirect fluorescent antibody test (IFAT). Journal of aquaculture Development 7(3): 53-61.

Ghasemzadeh, J. 1998. Phylogeny and systematics of Indo-Pacific mullets (Teleostei: Mugilidae) with special reference to the mullets of Australia. Ph.D. Thesis, Macquarie University, Sydney. 397p.

Ghasemzadeh, J. & Ivantsoff, W. 2004. Historical overview of mugilid systematics, with description of Paramugil (Teleostei: Mugiliformes: Mugilidae), new genus. Aqua, International Journal of Ichthyology 81: 9-22.

Ghelichi, A.; Ahmadi, M.R.; Oryan, S. & Jorjani, S. 2004. The efficacy of hypaphysial, LRH- A2 and HCG hormones in plasma steroid levels in captive grey mullet (Mugil cephalus L.). Journal of Marine Sciences and Technology 3(2-3): 61-68.

Ghelichi, A.; Akrami, R.; Bandani, G.A. & Jorjani, S. 2011. Comparison of oocyte development of common carp (Cyprinus carpio) and grey mullet (Mugil cephalus). Journal of Biology Science 4(15): 69-81.

Ghelichi, A.; Hajimoradlou, A.A. & Jorjani, S. 2007. 17α hydroxyprogesterone profiles during induced spawning of the grey mullet (Mugil cephalus). Journal of Agricultural Sciences and Natural Resources 14(2): 88-97. (In Farsi)

Ghelichi, A. & Jorjani, S. 2004. Sex steroid profiles and ovarian development during induced spawning of grey mullet (Mugil cephalus) by CPH and hCG. Biology in Asia InternationalConference, Singapore, 7-10 December 2004 (abstract).

Ghelichi, A.; Oryan, S.; Ahmadi, M.R.; Kazemi, R.A. & Halajian, A. 2004. The histological study of ovarian development of grey mullet (Mugil cephalus) in Gomishan fish and shrimp rearing center. Journal of Agricultural Sciences and Natural Resources 10(4): 115-124. (In Farsi)

Ghelichpour, M.; Shabanpour, B. & Hoseini, S. M. 2012. Comparative study on the effect of different cooking methods on physicochemical properties and color characteristics of golden grey mullet (Liza aurata) fillets. Latin American Applied Research 42: 155-159.

Ghiasi, M.; Binaii, M.; Ghasemi, M.; Fazli, H. & Zorriehzahra, M.J. 2016. Haemato-biochemical disorders associated with nodavirus like-agent in adult leaping mullet Liza saliens (Risso, 1810) in the Caspian Sea. VirusDisease 27(1): 12-18.

Ghodsi, Z.; Shabani, A. & Shabanpour, B. 2011. Genetic diversity of Liza aurata (Risso, 1801) in the coastal regions of Golestan Province, using microsatellite markers. Journal of Taxonomy and Biosystematics 3(6): 35-45.

Ghodsi, Z.; Shabani, A. & Shabanpour, B. 2013. Genetic diversity of Liza aurata (Risso, 1801) in the coastal regions of Mazandaran Province, using microsatellite markers. Modern Genetics Journal 8(1): 29-36.

Ghorbani Ranjbary, A.; Ghorbani Ranjbary, N.; Ghorbani Ranjbary, Z.; Marhamtizadeh, M.H. & Cheraghi, P. 2013. Determination of lead, mercury and cadmium concentrations in different organs of Barbus grypus and Liza abu of Karoon River in 2011. Journal of Food Hygiene 2(4)(8): 53-60.

Golchin Manshadi, A.R.; Masoumian, M.; Jalali Jafari, B. & Barzegar Dowlatabadi, M. 2012. Protozoan and myozoan infections in some fishes of Parishan Lake. Asian Journal of Animal and Veterinary Advances 7(9): 842-850.

Haghparast, M.M.; Hosseini, S. & Sajedi, A. 2015. Species characteristics of the mullet (Mugilidae). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Halstead, B.W. 1967-1970. Poisonous and Venomous Marine Animals of the World. Volume Two - Vertebrates. 1070p., Volume Three - Vertebrates continued. 1006p. United States Government Printing Office, Washington, D.C.

Halstead, B.W. 1978. Poisonous and Venomous Marine Animals of the World (Revised Edition). The Darwin Press, Princeton, New Jersey. 1043p.

Hantoush, A.A.; Al-Saad, H.T. & Abdul-Hussain, E.A. 1999. Seasonal variations of some biochemical aspects of the muscles of some freshwater and marine fishes from Shatt Al-Arab River and northwest Persian Gulf. Marina Mesopotamica 14(2): 427-453. (In Arabic)

Hashemi, S.; Eskandary, Gh.; Ansary, H. & Yooneszadeh, M. 2011. Stock assessment and production of fish species in the Shadegan Wetland, Iran. World Journal of Fish and Marine Sciences 3(6): 502-508.

Hashemi, S.A.; Ghorbani, R.; Kymaram, F.; Hossini, S.A.; Eskandari, Gh. & Hedayati, A. 2014. Estimation of fish composition and catchability coefficient of gillnet in the Shadegan Wetland. Iranian Journal of Ichthyology 1(1): 51-60.

Hashemi, S.A.; Ghorbani, R.; Kymaram, F.; Hossini, S. A.; Eskandari, G. & Hedayati, A. 2015. Fish species composition, distribution and abundance in Shadegan Wetland. Fisheries and Aquaculture Journal 6(2): 128.

Hashemi, S.A.R. 2010. Study of some biology characteristics of fish main species in the Shadegan wetland. National Wildlife Conference, Azad University, Ahvaz, March 2010. 13p. (In Farsi)

Hashemi, S.A.R. & Ansary, H. 2012. Biomass and production of fish species in the Shadegan Wetland, Iran. Global Veterinaria 9(2): 123-128.

Hashemi, S.A.R.; Eskandary, Gh. & Ansary, H. 2010a. Study of catch and biomass of fish species in the Shadegan wetland. National Wildlife Conference, Azad University, Ahvaz, March 2010. 11p. (In Farsi)

Hashemi, S.A.R.; Eskandary, Gh. & Ansary, H. 2010b. Study of catch and biomass of fish species in the Shadegan wetland. Wetland 1(4): 3-9.

Heckel, J.J. 1843. Ichthyologie.In: Russegger, J. Reisen in Europa, Asien und Afrika, mit besonderer Rücksicht auf die naturwissenschaftlichen Verhältnisse der betreffenden Länder, unternommen in den Jahren 1835 bis 1841 von Joseph Russegger. Schweitzerbart'sche Verlagsbuchhandlung, Stuttgart 1(2): 991-1099.

Hedayatifard, M.; Moeini, S.; Keyvan, A. & Yousefian, M. 2002. Quantitative and qualitative identification of fatty acids in muscle of golden mullet (Liza aurata). Iranian Journal of Marine Sciences 1(2): 73-78. (In Farsi)

Hedayatifard, M. & Nemati, S. 2009. Changes of roe fatty acids of kutum Rutilus frisii kutum and golden muller Liza aurata affected by salting. Journal of Fisheries 3(2): 1-10.

Hedayatifard, M. & Yousefian, M. 2009a. Composition of fatty acid in golden mullet (Liza aurata) fillet affected by dry-salting. The 6th Scientific Conference of Fisheries Resources, 3-4 March 2009, Basrah, p. 106 (abstract).

Hedayatifard, M. & Yousefian, M. 2009b. The fatty acid composition of golden mullet fillet Liza aurata as affected by dry-salting. Journal of Fisheries and Aquatic Sciences 5(3): 208-215.

Heidari, B.; Shabanipour, N.; Yavari, L. & Totani, M. M. 2009. Histomorphological investigation of Liza aurata (Risso, 1810) (Mugilidae) ovary in the late oogenesis in the Caspian Sea. Iranian Journal of Fisheries Sciences 8(1): 25-36.

Hoda, S.M.S. 1978. On some post-larval fishes from Iraqi waters. Biologia, Lahore 24(2): 223-235.

Holčík, J. & Oláh, J. 1992. Fish, fisheries and water quality in Anzali Lagoon and its watershed. Report prepared for the project - Anzali Lagoon productivity and fish stock investigations. Food and Agriculture Organization, Rome, FI:UNDP/IRA/88/001 Field Document 2: 109p.

Hora, S.L. & Misra, K.S. 1943. On a small collection of fish from Iraq. Journal of the Asiatic Society of Bengal, Science 9(1): 1-15.

Hosseini, S. M., Mirghaffari, N., Sufiani, N. M., Hosseini, S. V. & Ghasemi, A. F. 2013. Risk assessment of the total mercury in golden gray mullet (Liza aurata) from Caspian Sea. International Journal of Aquatic Biology 1(6): 258-265.

Hosseini Aghuzbeni, S.H. & Hajirezaee, S. 2015. Effects of co-culturing of western white shrimp, Litopenaeus vannamei and grey mullet, Mugil cephalus on diversity and incidence of Vibrio bacteria from shrimp ponds. Journal of Aquaculture Development 9(1): 25-31.

Hosseini Aghuzbeni, S.H.; Hajirezaee, S.; Matinfar, A.; Khara, H. & Ghobadi, M. 2017. A preliminary sudy on polyculture of western white shrimp (Litopenaeus vannamei) with mullet (Mugil cephalus): an assessment of water quality, growth parameters, feed intake efficiency and survival. Journal of Applied Animal Research 45(1): 247-251.

Hosseinie, S.V.; Rezaee, M.; Sahari, M.A. & Hosseini, H. 2004. The effect of ice storage on lipid quality and sensory evaluation of golden muillet (Liza aurata). Iranian Journal of Marine Sciences 3(1): 31-40. In Farsi.

Hussain, N.A. & Ali, T.S. 2006. Trophic nature and feeding relationships among Al Hammer marsh fishes, southern Iraq. Marsh Bulletin 1(1): 9-18.

Hussain, N.A.; Ali, T.S. & Saud, K.D. 1989. Seasonal fluctuations and composition of fish assemblage in the Shatt Al-Arab at Basrah, Iraq. Journal of Biological Sciences Research, Baghdad 20(1): 139-150.

Hussain, N.A.; Al-Saboonchi, A.A. & Hamza, H.A. 1987. Aspects of abundance and the food habit of Liza abu (Heckel) in the Karkheh River (Arabistan). Journal of Biological Sciences Research, Baghdad 18(1): 145-161.

Hussain, N.A., Mohamed, A-R.M.; Al Noor, S.A.; Coad, B.; Mutlak, F.M.; Ibrahem, M.; Al-Sudani, I.M.; Mojer, A.M.; Toman, A.J. & Abdad, M.A. 2006. Species composition, ecological indices, length frequencies and food habits of fish assemblages of the restored southern Iraqi marshes. Report by the University of Basrah, Basrah, Iraq. 114p.

Hussain, N.A. & Naama, A.K. 1989. Survey of fish fauna of Khor Al-Zubair, north-west Persian Gulf. Marina Mesopotamica 4(2): 161-197.

Hussain, N.A.; Younis, K.H. & Yousif, U.H. 1997. The composition of small fish assemblages in the River Shatt Al-Arab near Basrah (Iraq). Acta Hydrobiologica 39(1/2): 29-37.

Hussain, N.A.; Younis, K.H. & Yousif, U.H. 2001. Evaluation of environmental degradation in the fish assemblage of Shatt Al-Arab River. Pakistan Journal of Zoology 33(2): 93-98.

Ingham, S.E. No date. The biology and taxonomy of the Mugilidae. Zoology Library, British Museum (Natural History), London. 276p.

Islam, A.K.M.S. & Al-Nasiri, S.K. 1978. Some morphometric studies on khishni (Liza abu) from Basrah, Iraq. Zanco A4/3: 129-140.

Islam, A.K.M.S. & Khalaf, A.N. 1983. Diel patterns of feeding of khishni Liza abu (Heckel) in Rashdiyah Reservoir in Baghdad, Iraq. Indian Journal of Fisheries 29(1&2)(1982): 223-228.

Islam, A.K.M.S.; Khalaf, A.N.; Jafery, A.R. & Sadek, S. E. 1981. Seasonal patterns of feeding of khishni, Liza abu (Heckel) in Rashdiyah Reservoir, Baghdad, Iraq. Bangladesh Journal of Zoology 9(2): 151-158.

IUCN. 2015. The IUCN Red List of Threatened Species. International Union for the Conservation of Nature, Gland, Switzerland (http://www.iucnredlist.org/).

Ivanov, V.P. & Katunin, D. N. 2001. Report on the results of the Caspian Marine Expedition. Caspian egional Thematic Center for Management of Bioresources (CRTC MB), Caspian Environmental Programme, Baku, Azerbaijan. 2 volumes.

Jahanbakhshi, A. 2016. Acute toxicity test of dichromate potassium (K2Cr2O7) in grey mullet (Mugil cephalus). 7th Euro-Global Summit on Toxicology and Applied Pharmacology, 24-26 October 2016, Rome (abstract).

Jawad, L.A. & Öktener, A. 2007. Incidence of lordosis in the freshwater mullet, Liza abu (Heckel, 1843) collected from Atatürk Dam Lake, Turkey. Anales de Biologia 29: 105-113.

Jawad L.A.; Sadighzadeh Z. & Valinassab, T. 2010. Malformation of the caudal fin in the freshwater mullet, Liza abu (Actinopterygii: Mugilidae) collected from Karkhe River Iran. Anales de Biologia 32: 11-14.

Jawad, L.A.J. 2004. Asymmetry analysis in the mullet, Liza abu collected from Shatt al-Arab River, Basrah, Iraq. Bollettino del Museo Regionale di Scienze Naturali di Torino 21(1): 145-150.

Jolodar, M.N. & Abdoli, A. 2004. Fish Species Atlas of South Caspian Sea Basin (Iranian Waters). Iranian Fisheries Research Organization, Teheran. 110p. (In Farsi)

Jouladeh-Roudbar, A.; Vatandoust, S.; Eagderi, S.; Jafari-Kenari, S. & Mosuavi-Sabet, H. 2015. Freshwater fishes of Iran; an updated checklist. AACL Bioflux 8(6): 855-909.

Kakoolaki, S.; Akbary, P.; Zorriehzahra, M.J.; Salehi, H.; Sepahdari, A.; Afsharnasab, M.; Mehrabi, M.R. & Jadgal, S. 2016. Camellia sinensis supplemented diet enhances the innate non-specific responses, haematological parameters and growth performance in Mugil cephalus against Photobacterium damselae. Fish and Shellfish Immunology 57: 379-385.

Kalani, N.; Riazi, B.; Karbassi, A.R. & Moattar, F. 2014. Heavy metal concentration in Liza saliens muscle and human health risk estimates from fish consumption in Gomishan International Wetland, Iran. Journal of Aquatic Animals and Fisheries 5(17): 65-79.

Kalbassi, M.R.; Abdollahzadeh, E. & Salari-Joo, H. 2013. A review on aquaculture development in Iran. Ecopersia 1(2): 159-178.

Kamran, A. 2006. Survey on bony fish catches in Gorgan Gulf. Iranian Fisheries Research Organization, Agriculture Research and Education Organization, Ministry of Jihad-e-Agriculture. http://en.ifro.ir (abstract).

Kardel, F.; Mirzapour, F.; Omidzahir, S. & Akhoundian, M. 2016. Heavy metal concentrations in gill and liver tissues of Rutilus kutum and Chelon aurata in the coast of Babolsar, southern Caspian Sea. International Journal of Aquatic Biology 4(2): 81-88.

Karimpour, M. 1998. The ichthyofauna of Anzali Lagoon. Iranian Journal of Fisheries Sciences 7(2): 83-94, 9. (In Farsi)

Kashi, M.; Hashemi, S. & Al-Mukhtar, M. 2009. Some biological aspects of keeled mullet, Liza klunzingeri in the east coastal of Khuzestan, north-west of the Gulf. The 6th Scientific Conference of Fisheries Resources, 3-4 March 2009, Basrah, p. 82 (abstract).

Kashi, M.T.; Hashemi Seyed, A.R. & Safikhani, H. 2009. A study of some characteristics of Klunzingeri mullet (Liza klunzingeri) in coastal waters of Khuzestan. Journal of Fisheries 2(4): 23-30. (In Farsi)

Keivany, Y. & Zamani-Faradonbe, M. 2017. Length-weight and length-length relationships for seven fish species from the Zohreh River, Iran. Journal of Applied Ichthyology DOI: 10.1111/jai.13285.

Kennedy, W.P. 1937. Some additions to the fauna of Iraq. Journal of the Bombay Natural History Society 39(4): 745-749.

Khalaf, A.N.; Al-Yamour, K.Y.; Allouse, S.B.; Al-Jafary, A. & Sadek, S.E. 1986. Age, growth, length-weight relationships, and distribution of khishni Liza abu (Heckel) (Mugilidae) in a polluted habitat. Journal of Biological Sciences Research, Baghdad 17(2): 63-81.

Khalaf, K.T. 1961. The marine and freshwater fishes of Iraq. Ar-Rabitta Press, Baghdad. 164p.

Khayami, H.; Zolgharnein, H.; Salamat, N. & Movahedinia, A. 2015. Morphological variability of Liza klunzingeri (Day, 1888) from Bandar Abbas Port and Qeshm Island in northeastern Persian Gulf. Journal of Fisheries and Aquatic Sciences 10(3): 191-198.

Khayyami, H.; Movahedinia, A.; Zolgharnein, H. & Salamat, N. 2014a. Length-weight relationship of Mugil cephalus (Linnaeus, 1758) from Bandar Abbas Port and Qeshm Island in northeastern Persian Gulf. Advances in Applied Science Research 5(5): 19-23.

Khayyami, H.; Movahedinia, A.; Zolgharnein, H. & Salamat, N. 2014b. Morphological variability of Liza aurata (Risso, 1810), along the southern Caspian Sea. Journal of Basic and Applied Biology 67: 100-107.

Khayyami, H.; Movahedinia, A.; Zolgharnein, H. & Salamat, N. 2014c. Morphological variability of Liza abu (Heckel, 1843) from northwestern part of Persian Gulf. World Journal of Fish and Marine Sciences 6(5): 386-394.

Khayyami, H.; Zolgharnein, H.; Salamat, N. & Movahedinia, A. 2015a. Morphological variability of Mugil cephalus (Linnaeus, 1758) from Bandar Abbas Port and Qeshm Island in northeastern Persian Gulf. International Journal of Pure and Applied Zoology 3(2): 103-112.

Khayyami, H.; Zolgharnein, H.; Salamat, N. & Movahedinia, A. 2015b. Anatomy and histology of the stomach and pyloric caeca in Mugilidae, Liza aurata (Risso, 1810), L. abu (Heckel, 1843) and Mugil cephalus (Linnaeus, 1758). Journal of the Persian Gulf (Marine Science) 6(19): 59-66.

Khodabandeh, S.; Shahriari Moghadam, M. & Abtahi, B. 2007. The first report of Mugil auratus juvenile’s contamination by Myxobolus spp. in the Caspian Sea. Comparative Biochemistry and Physiology 146(4)(Supplement): S185.

Khodabandeh, S.; Shahriari Moghadam, M. & Abtahi, B. 2009. Changes in chloride cell abundance, Na+, K+-ATPase immunolocalization and activity in the gills of golden grey mullet, Liza aurata, fry during adaptation to different salinities. Cell Journal 11(1): 49-54.

Khoroshko, A.I. 1982. Population abundance and structure in the long-finned mullet (genus Liza, Mugilidae) during acclimation in the Caspian Sea. Journal of Ichthyology 22(6): 62-69.

Khoshnood, Z. 2014. Identification and study of fish species in Karkheh River (Iran). Transylvanian Review of Systematical and Ecological Research 16(2): 97-106.

Kiabi, B.H.; Abdoli, A. & Naderi, M. 1999. Status of the fish fauna in the South Caspian Basin of Iran. Zoology in the Middle East 18: 57-65.

Klinkhardt, M.; Tesche, M. & Greven, H. 1995. Database of Fish Chromosomes. Westarp Wissenschaften, Magdeburg. 237p.

Kohestan-Eskandari, S., AnvariFar, H. & Mousavi-Sabet, H. 2013. Detection of morphometric differentiation of Liza aurata (Pisces: Mugilidae) in southeastern of the Caspian Sea, Iran. Our Nature 11(2): 126-137.

Koshafar, A. & Velayatzadeh, M. 2015. Comparison of bioaccumulation of heavy metals in muscle of two species Liza abu and Acanthopagrus latus from Bahmanshir River in summer. Journal of Wetland Ecobiology 6(4): 59-72. (In Farsi)

Kuronoma, K. & Abe, Y. 1986. Fishes of the Persian Gulf. Kuwait Institute for Scientific Research, State of Kuwait. 357p.

Maghtouie, A.H.; Neissi, G.; Nasseri, S.; Gholaminezhad, E.; Shalamzari, N. & Nikpour, Y. 2011. Determination of mercury in mullet fish (Liza abu) from Arvand River, Iran. World Journal of Fish and Marine Sciences 3(6): 514-517.

Mahdi, A.A.; Faddagh, M.S.; Tuman, A.J. & Abdullah, T.A. 2005. Biochemical composition and caloric value of six freshwater fish species from the southern Iraqi marshes. First Scientific Conference on the Rehabilitation of the Southern Iraq Marshes, 11-12 April 2005, Marine Science Centre, University of Basrah (abstract).

Mahdi, N. 1962. Fishes of Iraq. Ministry of Education, Baghdad. 82p.

Malakootian, M.; Mortazavi, M.S. & Ahmadi, A. 2016. Heavy metals bioaccumulation in fish of southern Iran and risk assessment of fish consumption. Environmental Health Engineering and Management Journal 3(2): 61-68.

Mandych, A.F. 1995. Enclosed seas and large lakes of eastern Europe and middle Asia. SPB Academic Publishing, Amsterdam. 273p.

Marammazi, J.G. 1994. Preliminary ecological investigation of six fish species in Zohre River. Iranian Fisheries Scientific Journal 3(2): 57-70, 6. (In Farsi)

Marti, V.Yu. 1940. Akklimatizatsiya kefali v Kaspiiskom more [Acclimatization of the grey mullet in the Caspian Sea]. Priroda 1940(1): 88-90.

Marti, V.Yu. 1941. Novoe ob akklimatizatsii kefali v Kaspiiskom more [New data on the acclimatization of the grey mullet in the Caspian Sea]. Priroda 1941(3): 85-86.

Mashian Moradi, A. & Golshani, R. 2013. Determination of concentration of organophosphate pesticides (diazinon – malathion– azinfos methyl) in mullet (Liza aurata) muscle. International Journal of Marine Science and Engineering 2(4): 255-258.

Mashinchianmoradi, A. & Pirouznezhad, M. 2013. Concentration of heavy metals in liver and muscle tissue of Liza aurata in estuaries of the rivers Babolroud, Tajan and Gorganroud. International Journal of Marine Science and Engineering 3(1): 7-12.

Masroor, R.; Ramezani Rad, O.; Mazaheri, Y.; Khazaeel, K. & Pourmehdi Broujeni, M. 2015. Morphology and biometry of the gray mullet (Liza abu: Mugilidae). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Massoud, J.; Jalali, H. & Reza, M. 1981. Studies on trematodes of the family Heterophyidae (Odhner, 1914) in Iran: 1. Preliminary epidemiological surveys in man and carnivores in Khuzestan. Journal of Helminthology 55(4): 255-260.

Menon, A.G.K. & Yazdani, G.M. 1968. Catalogue of type-specimens in the Zoological Survey of India. Part 2.-Fishes. Records of the Zoological Survey of India 61(1 & 2): 1-190.

Mhaisen, F.T. & Al-Jaffery, A.R. 1989. Determination of age and growth of the mugilid fish Liza abu (Heckel) inhabiting Babylon Fish Farm, Hilla, Iraq. Journal of Biological Sciences Research, Baghdad 20(3): 547-554.

Mhaisen, F.T. & Yousif, U.H. 1988. Reproductive biology of the mugilid fish Liza abu (Heckel) from Mehaijeran Creek, Basrah. Basrah Journal of Agricultural Science 1: 48- 55.

Mhaisen, F.T. & Yousif, U.H. 1989. Age and growth of the mugilid fish Liza abu (Heckel) from Mehaijeran Creek, Basrah, Iraq. Bulletin of the Iraq Natural History Museum 8(2): 147-155.

Mikaili, P. & Shayegh, J. 2011. An etymological review on fish common and scientific names in the Euphrates and Tigris. Research Journal of Fisheries and Hydrobiology 6(4): 412-423.

Miller, P.J. 2003. The Freshwater Fishes of Europe. Volume 8/I. Mugilidae, Atherinidae, Atherinopsidae, Blenniidae, Odontobutidae, Gobiidae 1. AULA-Verlag, Wiebelsheim. 404p.

Mir Hashemi Rostami, S.A., Amini, K.; Khani, F. & Kolangi Miandare, H. 2016. Grey mullet (Mugil cephalus) rearing normative as a new species in Gomishan, Golestan Province. Environmental Resources Research 4(1): 47-55.

Mirhashemi Rostami, S.A.; Amini, K.; Jorjani, M.; Piri, H.; Hami Tabari, A.; Iri, Y.; Shafe, A.A.Gh.; Saghali, M. & Poursoufi, T. 2008. Mortality factors in artificially reproduced grey mullet (Mugil cephalus L.) fries. Iranian Scientific Fisheries Journal 17(3): 153-160. (In Farsi)

Mirhashemi Rostamy (sic), S.A.; Joorjani, M.; Amini, K.; Shafaee, A. & Ghazel, H.G. 2006. An investigation on artificial reproduction of Mugil cephalus. Iranian Journal of Fisheries Sciences 14(4): 181-196. (In Farsi)

Moghainemi, R.S. & Abbasi, S. 1992. Parasitic fauna of economically important fishes from Hoor-Elazim Marsh. Iranian Fisheries Research and Training Organization Publication, Tehran. 107 pp. In Farsi.

Moghim, M.; Ghani Nezhad, G. & Fazli, F. 1994. Stock assessment of the Caspian Sea bony fishes. Iranian Fisheries Journal 3(1): 35-52, 4. (In Farsi)

Mohamadi, M.; Bishkoul, G.R.; Rastiannasab, A.; Khara, H. & Hut, N. 2014. Physiological indicators of salinity stress in the grey muller, Mugil cephalus, Linnaeus, 1758 juveniles. Comparative Clinical Pathology 23(5): 1453-1456.

Mohamed, A-R.M. 2014. Stock assessment of freshwater mullet, Liza abu (Heckel, 1843) populations in three restored southern marshes, Iraq. Croatian Journal of Fisheries 72: 48-54.

Mohamed, A-R.M., Abood, A.N. & Hussein, S.A. 2016. Comparative taxonomical study of four mullet species (Mugiliformes: Mugilidae) from Iraqi marine waters, Persian Gulf. Basrah Journal of Agricultural Sciences 29(2): 11-23.

Mohamed, A-R.M.; Alokailee, M. & Sumaea, M.A. 2013. Abundance and distribution of Liza abu eggs and larvae in north part of Shatt Al-Arab River. The 1st Scientific Conference of Science College, University of Karbala, Karbala, Iraq, January 2013 (abstract).

Mohamed, A.R.M., Hussein, S.A. & Mutlak, F.M. 2013. Stock assessment of greenback mullet, Liza subviridis in East Hammar Marsh, Iraq. Basrah Journal of Agricultural Sciences 26(1): 315-327. (In Arabic)

Mohamed, A.R.M.; Hussein, S.A. & Saleh, J.H. 1998. The biology of green back grey mullet Liza subviridis in the northwest Persian Gulf. Marina Mesopotamia 13(2): 375-386.

Mohammadshahali, S.; Masoudifard, M.; Vajhi, A.; Moghim, M.; Zehtabvar, O. & Rostami, A.M. 2014. Ultrasonography of the striped mullet (Mugil cephalus Linnaeus, 1758) for sex determination. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Mokhayer, B. 1973. Study on the parasitism of Mugil auratus Risso (sic) from the out (sic, south) Caspian Sea. Rivista italiana di Piscicoltura e Ittiopatologia 8(2): 53-54.

Mokhayer, B. 1976. Fish diseases in Iran. Rivista italiana di Piscicoltura e Ittiopatologia 11(4): 123-128.

Mokhayer, B. 1981. Infection of Karoun River and Arvandroud mullet with Contracaecum larvae. Journal of the Veterinary Faculty of the University of Tehran 37(1): 91-102. (In Farsi)

Moradinasab, A.; Babaei, N.; Parsa, M.; Raeisi, H.; Daliri, M.; Rahnama, B.; Nekuro, A. & Vesaghi, M.J. 2015. By-catch composition of kilka fishes (Clupeonella) lantern net in the southwest of the Caspian Sea (Bandar Anzali fishing grounds). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Moradinasab, Gh.; Raeisi, H.; Paighambari, S.Y.; Ghorbani, R. & Bibak, Z. 2012. Length-weight relationships, relative condition factor and relative weight of three fish species from beach seine fishing grounds in Iranian coastal waters of Caspian Sea. Scientific Research and Essays 7(18): 1809-1812.

Mortezaei, S.; Mobedi, I. & Farahnak, A. 2000. Infection of some species of fresh water fishes to parasitic worms in Khouzestan Province, Iran. Iranian Journal of Fisheries Sciences 9(1): 25-36. (In Farsi)

Mostafavi, H. & Abdoli, A. 2003. A study of fish species diversity in Talar River in Mazandaran. Environmental Sciences 1(1): 20-29. (In Farsi)

Movchan, Yu.V. 1988. Fauna of Ukraine. Fishes. Loaches, catfishes, channel catfishes, freshwater eels, conger eels, garfishes, codfishes, sticklebacks, seahorses, mosquitofishes, John dories, barracudas, grey mullets, silversides, lings. Naukova Dumka Publishing House, Kiev 8(3): 367 p.

Muhsin, K.A. 1988. Annual cycle and body composition of female Liza subviridis from Khor Al-Zubair, north west Persian Gulf. Marina Mesopotamica 3(2): 125-137.

Naama, A.K., Ahmed, H.A. & Al-Adhub, A.H.Y. 1986. Aspects of reproduction of the mullet Liza abu (Heckel)(Pisces, Mugilidae) in Al-Hammar Marsh, Iraq. Cybium 10(1): 47-55.

Naama, A.K. & Muhsen, K.A. 1986. Feeding periodicites (sic) of the mugilid Liza abu (Heckel) and cyprinid Carasobarbus luteus (Heckel) from Al-Hammar Marsh, southern Iraq. Indian Journal of Fisheries 33(3): 347-350.

Naem, S.; Mobedi, I.; Khomirani, R. & Abolghasemi, S. J. 2002. A survey on gill parasites in cultured and wild fishes of western branch of Sepid-Rood river in Gilan province, Iran and introducing new species. Iranian Journal of Veterinary Research, Shiraz University 3(2): 118-128. (In Farsi)

Najafpour, N. 1997. Identification of some freshwater fishes of Khuzestan Province. Fisheries Research Centre of Khuzestan Province, Iranian Fisheries Research and Training Organisation, Tehran. 96p. (In Farsi)

Naseri, F.; Abedian Amiri, A.; Hosseini, M.; Heidarzadeh, M.; Zahedi, R. & Arabzadeh, P. 2014. Molecular and phenotypic characterization of Photobacterium damselae, a pathogen of the grey mullet Mugil cephalus, in Iran. International of Advanced Life Sciences 7(3): 452-462.

Nasir, N.A. & Na’ama, A.K. 1988. Liza abu Heckel, 1843 (Pisces, Mugilidae): a new record from Khor Al-Zubair, North-west of the Persian Gulf. Mahasagar 21(2): 113-115.

Nasrollahzadeh Saravi, H.; Pourgholam, R.; Unesipour, H. & Makhlough, A. 2012. Study of polyaromatic hydrocarbons (16PAHs) at the sediments and edible tissue of Liza saliens and Rutilus frisii kutum of the Caspian Sea. Journal of Mazandaran University of Medical Sciences 22(94): 79-89.

Nejad Shamoushaki M.M.; Dordee, K.; Rezaei Shirazi, A. & Yahiaee, M. 2013. Study of trend process of (Rutilus frisii kutum), (Liza spp.) and (Cyprinus carpio) in provinces of Golestan, Mazandaran and Guilan (2009-2011). Journal of Aquatic Animals and Fisheries 4(15): 27-38.

Nelson, J.S.; Grande, T.C. & Wilson, M.V.H. 2016. Fishes of the World. Fifth Edition. John Wiley & Sons, Hoboken. 707p.

Nematzadeh, M.; Rezvani, S.; Khalesi, M.K.; Laloei, F. & Fahim, A. 2013. A phylogeny analysis on six mullet species (Teleosti: Mugillidae) using PCR-sequencing method. Iranian Journal of Fisheries Sciences 12(3): 669- 679.

Nematzadeh, M.; Rezvani Gillkolaei, S.; Khalesi, M.K., Laloei, F. 2013. Molecular phylogeny of mullets (Teleostei: Mugilidae) in Iran based on mitochondrial DNA. Biochemical Genetics 51(3-4): 334-340.

Nezami, S.A.; Savari, A.; Sakari, M. & Alizadeh, M. 2000. National Report of Biodiversity in Caspian Coastal Zone. Research Department, Gilan Provincial Office, Department of the Environment Conservation, Iran (TACIS (Technical Assistance to the Commonwealth of Independent States, European Union), Caspian Environmental Programme). 97p.

Noori, M. & Lalouie, F. 1994. Studying Mugil sp. fishing period from the management point of view. Iranian Fisheries Scientific Journal 3(2): 71-77, 7. In Farsi.

Norouzi, M. & Bagheri, M. 2015. The chemical composition of golden grey mullet Liza aurata in southern Caspian Sea during sexual rest and sexual ripeness. AACL Bioflux 8(4): 517-525.

Norouzi, M. & Bagheri Tavani, M. 2016. The effects of breeding season and habitat on the biochemical composition and energy content of the Liza aurata fillet (Teleostei: Mugilidae). Iranian Journal of Ichthyology 3(4): 283-293.

Özdilek, Ş.Y. 2003. Occurrence of the abu mullet, Liza abu (Heckel, 1843) (Pisces, Mugilidae), in the Orontes River. Zoology in the Middle East 30: 111-113.

Papahn, F.; Rezaei, M.; Eskandari, Gh. & Rasekhi, A. 2013. The survey of fish population Hoor-Al Azim Marsh. Journal of Wetland Ecobiology 5(16): 33-40. (In Farsi)

Patimar R. 2008. Some biological aspects of the sharpnose mullet Liza saliens (Risso, 1810) in Gorgan Bay-Miankaleh Wildlife Refuge (the Southeast Caspian Sea). Turkish Journal of Fisheries and Aquatic Sciences 8(2): 225-232.

Pazira, A.; Abdoli, A.; Ghanbari, F. & Moghdani, S. 2016. Comparison of fish species diversity in Dalaki and Helleh Rivers of the Persis basin in Bushehr Province. Iranian Journal of Ichthyology 3(3): 222-228.

Pazooki, J.; Abtahi, B. & Rezaei, F. 2009. Determination of heavy metals (Cd, Cr) in the muscle and skin of Liza aurata from the Caspian Sea (Bandar Anzali). Environmental Sciences 7(1): 21-31.

Pietschmann, V. 1912. Eine neue Mugil-Art aus dem Schatt el Arab. Anzeiger der Akademie der Wissenschaften, Wien, Mathematische-naturwissen schaftliche Klasse 49(17): 268-270.

Pillay, S.R. 1972. A bibliography of the grey mullets Family Mugilidae. Food and Agriculture Organization Fisheries Technical Paper 19: 99 p.

Piri, I.; Nezhad, I.M. & Amirian, P. 2009. Investigation of fish species diversity in Haraz River. Proceedings of the 2009 International Conference on Chemical, Biological and Environmental Engineering (CBEE 2009), World Scientific Publishing, Singapore, p. 323-328.

Pourfaraj, V.; Karami, M.; Nezami, S.A.; Rafiee, G.R.; Khara, H. & Hamidoghli, A. 2013. Study of some biological features of mullets in Iranian coasts of the Caspian Sea. Journal of Utilization and Cultivation of Aquatics 1: 97-110. (In Farsi)

Rabbaniha, M., Abdolmaleki, D. & Khedmati, K. 2015. Hydrological parameters affected on Mugilidae caught fluctuation in the Caspian Sea – Bandar Anzali (2003-2012). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

RaLonde, R. & Walczak, P. 1972. A review of Shilot's bony fishing policy from 1929-1972. Report of the Fisheries Research Institute, Bandar Pahlavi. MS, 10 p.

Ramezani Rad, O.; Masroor, R.; Mazaheri, Y.; Khazaeel, K.; Mohammadian, T. & Ghadiri, A. 2015. Neuro cranial survey of gray mullet (Liza abu: Mugilidae). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Ramin, M. 1997. Indentification (sic) of Babolrud River's fish fauna. Iranian Fisheries Scientific Journal 6(3): 59-72, 8. (In Farsi)

Randall, J.E. 1995. Coastal Fishes of Oman. University of Hawai'i Press, Honolulu. 439p.

Rashid, K.H. 1994. Histological changes in the ovarian stages of Liza abu (Heckel). Marina Mesopotamica 8(1)(1993): 167-177.

Rasta, M.; Rahimibashar, M.R.; Nia, M.R.A. & Torabi, H. 2015. Reproductive biology of Chelon subviridis in north of Persian Gulf (Bushehr province). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Razavilar, V. & Tavakoli, H.R. 2006. A prevalence study of human toxigenic types of Clostridium botulinum (A, B, E) in some sea water fishes of northern (Rutilus frisii kutum and Mugil auratus Risso) and southern (Otolithes ruber and Stromateus niger) regions of Iran. Journal of Veterinary Research 61(1): 39-42. In Farsi.

Razivi, B.; RaLonde R. & Walczak, P. 1972. 1972 report on stock assessment and composition of the commercial bony fishes of the Southern Caspian Sea. Report of the Fisheries Research Institute, Bandar Pahlavi 32p.

Rezvani Gilkolaei, S.; Ghane, R.; Fazli, H.; Kamali, A.; Lalouie, F. & Taghavi, M.J. 2012. Genetic iversity of Liza saliens in the coastal waters of Guilan and Golestan province using the mitochondrial DNA sequencing (mtDNA). Journal of Marine Biology 4(14): 37-44.

Robins, C.R.; Bailey, R.M.; Bond, C.E.; Brooker, J.R.; Lachner, E.A; Lea, R.N. & Scott, W.B. 1991. World Fishes Important to North Americans Exclusive of Species from the Continental Waters of the United States and Canada. American Fisheries Society Special Publication 21: 243p.

Roshan Tabari, M. 1997. Hydrological and hydrobiological study in the Syiah-rud River. Iranian Fisheries Scientific Journal 6(2): 27-42 (In Farsi).

Saeedi, A.A.; Khoshbavr Rostami, H.A.; Kamgar, M.; Rezvani, Z.; Zorriehzahra, M.E.J. & Sharifpour, I. 2004. The comparison of some important hematological and serum biochemical parameters in healthy and diseased golden grey mullets (Liza auratus (sic)) fishes in southern coasts of Caspian Sea. Proceedings of the 4th International Iran and Russia Conference "Agriculture and Natural Resources", 8-10 September, Shahr-e Kord, Iran, p. 1518-1521.

Saeidi, Z.; Rezvani Gilkolaei, S.; Soltani, M. & Laloei, F. 2014. Population genetic studies of Liza aurata using D-Loop sequencing in the southeast and southwest coasts of the Caspian Sea. Iranian Journal of Fisheries Sciences 13(1): 216-227.

Saghali, M.; Baqraf, R.; Hosseini, S.A.; Nejatkhah Manavi, P. & Moghimi Tilemi, B. 2012. Heavy metals (Zn, Pb, Cd and Cr) in fish, water and sediments of southern Caspian Sea, Iran. The First International Conference on Larviculture in Iran and International Workshop on Replacement of Fish/Meal Oil with Plant Sources in Aquatic Feed, Iran-Larvi 2012, 11-12 December 2012, Karaj, Iran, p. 607-618.

Sahari, M.A.; Ahmadnia, A.; Barzegar, M. & Noorolahi, Z. 2014. Vitamin loss during frozen storage of Liza aurata (Risso, 1810), Cyprinus carpio L., 1758, Clupeonella cultriventris caspia (Nordmann, 1840), Rutilus frisii kutum (Kamenskii, 1901) and Sander lucioperca (L., 1758). Journal of Applied Ichthyology 30(2): 366-371.

Sakri, M. 1993. Shahrivar (September), the month of spawning and retainment for mullets. Abzeeyan, Tehran 4(7): 11. In Farsi.

Salehi, V.; Askari Hesni, M.; Teimori, A. & Lashkari, M. R. 2016. The sagittal otolith morphology of four selected mugilid species from Iranian waters of the Persian Gulf. International Journal of Aquatic Biology 4(5): 318-324.

Salim, S.M. 1962. Marsh Dwellers of the Euphrates Delta. London School of Economics, Monograph on Social Anthropology, 23, University of London, The Athlone Press. 157p.

Samadani, A.A. & Gharahveysi, S.A. 2010. The morphometric and polymorphism investigation of transferrin, total protein and hemoglobin of (Liza auratus) in the southern Casian Sea. Journal of Marine Biology 1(4): 85-94. In Farsi.

Sayari, H. & Rahmani, H. 2014. Morphological comparison of two sexes of garyz (Liza subviridis) in Persian Gulf, coastal area of Bandar Abbas. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Sayyadzadeh, G.; Esmaeili, H.R.; Ghasemian, S.; Mirghiyasi, S.; Parsi, B.; Zamanpoore, M. & Akhlaghi, M. 2016. Co-invasion of anchor worms Lernaea cyprinacea (Copepoda: Lernaeidae) in some freshwater fishes of the Kor River basin, southwest of Iran with some remarks on the ecological aspects of lernaeosis in the country. Iranian Journal of Fisheries Sciences 15(1): 369-389.

Shabanipour, N. 1995. Histomorphological study on the ovary of Mugil auratus (Risso). Iranian Fisheries Scientific Journal 4(2): 47-62, 6. (In Farsi)

Shabanipour, N. & Heidari, B. 2004. A histological study of the zona radiata during late oocyte developmental stages in the Caspian Sea mugilid Liza aurata (Risso 1810). Brazilian Journal of Morphological Sciences 21(4): 191-195.

Shadi, A.; Dehghan Mediseh, S.; Kouchanian, P. & Gandomi, Y. 2011. Length-weight relationships for 6 fish species from Khuzestan (north of Persian Gulf), Iran. World Journal of Fish and Marine Sciences 3(2): 129-131.

Shahriari Moghadam, M.; Abtahi, B.; Mosafer Khorjestan, S. & Bitaab, M.A. 2013. Salinity tolerance and gill histopathological alterations in Liza aurata Risso, 1810 (Actinopterygii: Mugilidae) fry. Italian Journal of Zoology 80(4): 503-509.

Shahriari Moghadam, M., Khodabandeh, S. & Ghanizadeh Kazerouni, E. 2006. Accumulation of mercury and its histopathological effect on kidney and gill of (Liza aurata). Journal of Marine Sciences and Technology 5(1-2): 19-27.

Sharifpour, I.; Yousefian, M. & Khalesi, M.K. 2003. Reproduction cycle and histology of the oocyte maturation in cultured grey mullet (Mugil cephalus). Iranian Journal of Marine Sciences 2(1): 23-34. (In Farsi)

Sharma, K.P. 1980. Further studies on the fish marketing conditions of Southern Iraq. The Arab Gulf, University of Basrah 2(1): 223-228.

Shayegi, M.; Khoobdel, M.; Bagheri, F.; Abtahi, M. & Zeraati, H. 2012. Organophosphorous residue in Liza aurata and Cyprinus carpio. Asian Pacific Journal of Tropical Biomedicine 2(7): 564-569.

Shokrzadeh Lamuki, M.; Saeedi Saravi, S.S. & Otadi, N. 2012. Evaluation of residues of D.D.T. and D.D.A. in fish collected from Caspian Sea, Iran. Iranian Journal of Toxicology 6(18): 704-708.

Shukolyukov, A. 1937. K voprosu ob akklimatizatsii cherno-morskikh ryb v Kaspiiskom more [Acclimatization of Black Sea fish in the Caspian Sea]. Rybnoe Khozyaistvo 6: 34-35.

Soltani, M. & Rahanandeh, M. 2001. Report of blout (sic) in Mugil capito in the Caspian Sea. Journal of the Faculty of Veterinary Medicine, University of Tehran 56(1): 106-107. (In Farsi)

Soltani, M.; Ghasemi, M.; Sharifrohani, M.; Sharifpour, I. & Zorrieh Zahra, J. 2010. Isolation and identification of betanodavirus causing mass mortalities in golden grey mullet (Liza auratus) in the Caspian Sea. Iranian Journal of Veterinary Research 4: 201–208.

Syzpuła, J.; Epler, P.; Bartel, R. & Szczerbowski, J.A. 2001. Age and growth of fish in lakes Tharthar, Razzazah and Habanniya. Archiwum Rybactwa Polskiego 9(supplement 1): 185-197.

TACIS (Technical Assistance to the Commonwealth of Independent States, European Union) & United Nations Development Programme. 2002. Transboundary diagnostic analysis for the Caspian Sea. Caspian Environment Programme, Baku, Azerbaijan. Volume 1: 26p., Volume 2: 128p., Volume 3: 13 p.

Talwar, P.K. & Kacker, R.K. 1984. Commercial Sea Fishes of India. Handbook No. 4, Zoological Survey of India, Calcutta. 997p.

Tehrani, M.G. 2006. Culturing of mullet fingerlings species Mugil cephalus L. Iranian Fisheries Research Organization, Agriculture Research and Education Organization, Ministry of Jihad-e-Agriculture, Sari. http://en.ifro.ir (abstract).

Teimori, A.; Esmaeili, H.R. & Gholamhosseini, A. 2010. The ichthyofauna of Kor and Helleh River basins in southwest of Iran with reference to taxonomic and zoogeographic features of native fishes. Iranian Journal of Animal Biosystematics 6(1): 1-8.

Thomson, J.M. 1963. Synopsis of biological data on the grey mullet Mugil cephalus Linnaeus 1758. Commonwealth Scientific and Industrial Research Organization (CSIRO), Melbourne, Fisheries and Oceanography, Fisheries Synopsis 1: 74p.

Thomson, J.M. 1997. The Mugilidae of the world. Memoirs of the Queensland Museum 41(3): 457-562.

Trewavas, E. & Ingham, S.E. 1972. A key to species of Mugilidae (Pisces) in the Northeastern Atlantic and Mediterranean, with explanatory notes. Journal of Zoology 167(1): 15-29.

Turan, C.; Ergüden, D.; Turan, F. & Gürlek, M. 2004. Genetic and morphologic structure of Liza abu (Heckel, 1843) populations from the rivers Orontes, Euphrates and Tigris. Turkish Journal of Veterinary and Animal Sciences 28: 729-734.

Ünlü, E.; Balci, K. & Meriç, N. 2000. Aspects of the biology of Liza abu (Mugilidae) in the Tigris River (Turkey). Cybium 24(1): 27-43.

Valinasab, T.; Seyfabadi, S.J.; Javadzadeh, N. & Safikhani, H. 2004. Fecundity of Klunzinger mullet, Liza klunzingeri, in coastal waters of Hendijan (Persian Gulf). Iranian Journal of Marine Sciences 3(1): 75-82. (In Farsi)

Valinassab, T.; Seyfabadi, S.J.; Javadzadeh, N. & Safikhani, H. 2007. Reproduction of Klunzinger mullet, Liza klunzingeri, in coastal waters of Khuzestan Province (Persian Gulf). Iranian Journal of Fisheries Sciences 6(2): 129-142.

van den Eelaart, A. 1954. Report to the Government of Iraq on the development of inland fisheries. Food and Agriculture Organization, Rome, EPTA Report 270: 1-42.

Vazirzadeh, A. & Ezhdehakoshpour, A. 2014. The effect of different hormonal treatments on the oocyte maturation in wild grey mullet (Mugil cephalus) collected from the Iranian coastal waters of the Oman Sea. Iranian Journal of Ichthyology 1(1): 17-22.

Velayatzadeh, M.; Askary Sari, A.A.; Beheshti, M. & Khodadadi, M. 2010. Study of the heavy metals concentration Cu and Zn in the muscle tissues of Liza abu in the Karoon, Bahmanshir and Dez rivers. Journal of Wetland Ecobiology 1(2): 59-69. (In Farsi)

Velayatzadeh, M.; Nildar, A. & Razavi, S.M.R. 2012. Effect of salt and deep freezing on drip and drip protein and TVB-N in four species fishes Liza aurata, Esox lucius, Cyprinus carpio and Rutilus frisii kutum from Caspian Sea. Journal of Aquatic Animals and Fisheries 3(11): 51-58.

Vladykov, V.D. 1964. Report to the Government of Iran on the inland fisheries, especially of the Caspian Sea with special reference to sturgeon. Food and Agriculture Organization, Rome, Report FAO/EPTA 1818: 51 pp.

Wahab, N.K. 1986. Ecology and biology of three species of mugilid fishes in Shatt Al-Basrah Canal. M.Sc. Thesis, University of Basrah. 155p. (In Arabic)

Wahab, N.K. 1999. Toxicity of four organophosphates insecticides on young Liza abu L. Marina Mesopotamica 14(1): 215-227. (In Arabic)

Whitfield, A.K.; Panfili, J. & Durand, J.-D. 2012. A global review of the cosmopolitan flathead mullet Mugil cephalus Linnaeus 1758 (Teleostei: Mugilidae), with emphasis on the biology, ecology and fisheries aspects of this apparent species complex. Reviews in Fish Biology and Fisheries 22(3): 641-681.

Xia, R.; Durand, J-D. & Fu, C. 2016. Multilocus resolution of Mugilidae phylogeny (Teleostei: Mugiliformes): implications for the family’s taxonomy. Molecular Phylogenetics and Evolution 96: 161-177.

Yeganeh, S.; Mojazi Amiri, B.; Yousefian, M. & Nematollahi, M.A. 2005. The influence of extenders on motility duration of gray mullet (Mugil cephalus) spermatozoa. Iranian Journal of Natural Resources 58(2): 383-393.

Yelghi, S.; Shirangi, S.A.; Ghorbani, R. & Khoshbavar Rostami, H.A. 2012. Annual cycle of ovarian development and sex hormones of grey mullet (Mugil cephalus) in captivity. Iranian Journal of Fisheries Sciences 11(3): 693-703.

Yesser, A.K.T.; Hinde, M.J. & Ahmed, H.A. 1999. Effects of starvation on proximate chemical composition of Liza abu (Heckel, 1843). Marina Mesopotamica 14(1): 11-17.

Yousefian, M.; Ghanei, M.; Pourgolam, M. & Rostami, H. K. 2009. Gonad development and hormonal induction in artificial propagation of grey mullet, Mugil Cephalus (sic) L. Research Journal of Fisheries and Hydrobiology 4(2): 35-40.

Yousefian, M.; Oryan, Sh.; Farokhi, F. & Asaeian, H. 2003. A study on oogenesis of Liza saliens. Iranian Journal of Fisheries Sciences 12(1): 131-152. (In Farsi)

Zandavar, H. & Norouzi, M. 2015. Diet of Liza aurata at southeast coast of Caspian Sea (Golestan Province). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015 (abstract).

Zare, M.; Abdoli, A.; Khoramian, S. & Borhani, M. 2014. Length-weight relationship and condition factor of beeyah (Mugilidae: Liza abu), in the Karkheh River, Andimeshk. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Zorriehzahra, J.; Ghiasi, M. & Binaii, M. 2014. The study on some hematological and biochemical parameters in golden grey mullet (Liza auratus) in southern Caspian Sea of Mazandaran Province following the occurrence of an emerged disease. Journal of Aquaculture Development 7(4): 25-33.

Zorriehzahra, M.E.J.; Ghasemi, M.; Ghiasi, M.; Haghighi Karsidani, S.; Bovo, G.; Nazari, A.; Adel, M.; Arizza, V. & Dhama, K. 2016. Isolation and confirmation of viral nervous necrosis (VNN) disease in golden grey mullet (Liza aurata) and leaping mullet (Liza saliens) in the Iranian waters of the Caspian Sea. Veterinary Microbiology 190: 27-37.

Zorriehzahra, M.J.; Nakai, T.; Gomez, D.K.; Sharifpour, I.; Shau-Chi, C.; Soltani, M.; Daud, H.H.M.; Rohani, M.S. & Saidi, AA.A. 2005. Mortality of wild golden grey mullet (Liza auratus) (sic) in Iranian waters of Caspian Sea, associated with viral nervous necrosis. Borok II, International Workshop, Invasions of Alien Species in Holarctic, 27 September-1 October 2005, Borok (abstract).

Zorriehzahra, M.J.; Nakai, T.; Sharifpour, I.; Kaw Gomez, D.; Shau-Chi, C.; Soltani, M.; Mohd Daud, H.M.; Sharif Rohani, M. & Saidi, A.A. 2005. Mortality of wild golden grey mullet (Liza auratus) (sic) in Iranian waters of the Caspian Sea, associated with viral nervous necrosis-like agent (2005). Iranian Journal of Fisheries Sciences 4(2): 43-58, 116-117.

Zorriehzahra, M.J.; Nazari, A.; Ghasemi, M.; Ghiasi, M.; Haghighi Karsidani, S.; Giuseppe, B. & Mohd Daud, H.H. 2014. Vacuolating encephalopathy and retinopathy associated with a nodavirus-like agent: a probable cause of mass mortality of wild Golden grey mullet (Liza aurata) and Sharpnose grey mullet (Liza saliens) in Iranian waters of the Caspian Sea. Virus Disease 25(4): 430-436.

Refbacks

  • There are currently no refbacks.